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Mark Jude Tramo. MI). PhD
Curriculum vitae
PART I: General Information
DATE PREPARED: June I 2010
Name
Mark Jude Tramo, M.D., Ph.D.
Office Address
1245 Wilshire Blvd, Suite 316
90017
Home Address
Place of Birth
Education
1978. B.A., Yale College
1982, M.D., Cornell University Medical College
1998, Ph.D. (Neurobiology), Harvard Graduate School of Arts & Sciences
Postdoctoral Training
1982-1983, Intern in Internal Medicine, LA County - USC Medical Center
1983-1986, Resident in Neurology. The New York Hospital - Cornell University Medical
Center
1985, Chief Resident. Neurology Ward. The New York Hospital
1986, Chief Resident. Neurology Ward. Memorial Hospital - Sloan-Kettering Cancer Center
1986-1987, Fellow (Cognitive Neuroscience), Dept. of Neurology, The New York Hospital - Cornell
University Medical Center and Memorial Hospital - Sloan-Kettering Cancer Center
Academic Appointments
1987-1988, Assistant Professor of Neurology, Cornell University Medical College
1988-1992. Assistant Professor of Medicine (Neurology) and of Psychiatry, Dartmouth Medical School
1992-1998. Instructor in Neurobiology, Harvard Medical School
1999-2007, Assistant Professor of Neurology, Harvard Medical School
2000-2005. Faculty. Harvard-M.I.T. Speech & Hearing Bioscience & Technology Graduate Studies Program
2000-2009. Board of Honors Tutors, Department of Psychology, Harvard University
2000-current, Faculty Fellow, Mind Brain & Behavior Interfaculty Initiative. Harvard University
2002-2009. Teaching Affiliate, Department of Biology (Neurobiology). Harvard University
2008-2009. Associate Professor of Neurology, Harvard Medical School
2009-current. Visiting Professor of Ethnomusicology. Herb Alpert School of Music at UCLA
Hospital and Affiliated Institution Appointments
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Mark Judc Tram°. April 2009
July 1987 — June 1988, Attending Neurologist. The New York Hospital. NY, NY
July 1988 — June 1991, Attending Neurologist, Dartmouth-Hitchcock Memorial Hospital, Hanover, NH
July 1988 - Sept 1991, Neurology Consultant, Franklin Regional Hospital, Franklin, NH
July 1992 —Dec 2009. Attending Neurologist. Massachusetts General Hospital. Boston. MA
July 1992 — June 2005, Neurology Consultant, McLean Hospital, Belmont, MA
July 1996 — Dec 2009. Research Associate. Eaton-Peabody Laboratory of Auditory Physiology. Massachusetts
Eye & Ear Infirmary, Boston, MA
July 1996 — current. Research Affiliate, Research Laboratory of Electronics, Department of Electrical
Engineering & Computer Science, Massachusetts Institute of Technology. Cambridge. MA
July 1999 — June 2005, Associate Neurologist, Affiliated Staff, Brigham & Women's Hospital, Boston, MA
Dec 2009 —current. Associate Neurologist. Affiliated Staff. Massachusetts General Hospital, Boston. MA
Dec 2009 — current. Medical Director, Neurological & Rehabilitation Services, and Staff Neurologist, Good
Samaritan Hospital. Los Angeles, CA
Other Professional Positions and Major Visiting Appointments
1998-current, Advisory Board, Songs of Love Foundation. New York. NY
2000-2005, Advisory Board, BioMusic Program, National Musical Arts. Washington, DC
2000. Consultant. Berkshire Hills Music Academy. South Hadley. MA
2002-current, Scientific Advisory Board, Science Museum of Minnesota. St. Paul, MN
2004-2007, Science Advisor, Morphonix, Inc., Sausalito. CA
2003-current. Founding Director, The Institute for Music & Brain Science, Boston. MA
2007-current, Advisory Board, National Center for Human Performance, Texas Medical Ctr. Houston. TX
Licensure and Certification
1983. Diplomate, National Board of Medical Examiners
1983-current, Drug Enforcement Administration Certification, U.S.A.
1983.1988, New York State Medical License
1988, Diplomate in Neurology. American Board of Psychiatry and Neurology
1988-1992, New Hampshire Medical License
1991-current, Massachusetts Medical License
1991-current, Massachusetts Drug Enforcement Administration Certification
21X19-current, California Medical License
Hospital and Health Care Organization Service Responsibilities
1986-1987, Fellow, then Attending, Stroke and Coma Team. The New York Hospital. NY. NY
1986-1988, Staff Physician, Medical and Neurological Services, Burke Rehabilitation Center, White Plains, NY
1987-1988. Attending, Neurology Consult Service. Behavioral Neurology Consult Service, Neurology Faculty
Practice Group, and Neurology Residents Clinic, The New York Hospital
1988-1991, Attending, Behavioral Neurology Consult Service and Neurology Residents Clinic, Mary Hitchcock
Memorial Hospital. Hanover, NH
1988-1991, Neurology Consultant, Franklin Regional Hospital. Franklin. NH
1992-2009, Attending, Neurology Inpatient Service, General Neurology Consult Inpatient Service, and
Cognitive/Behavioral Neurology Outpatient Unit, Massachusetts General Hospital, Boston. MA
1995-2005, Neurology Inpatient and Outpatient Consult Services, McLean Hospital, Belmont, MA
2009-current. Medical Director. Neurological & Rehabilitation Services, & Staff Neurologiet, Good Samaritan
Hospital, LA
Major Committee Assignments
1996-2000, Standing Committee on Neuroscience, Harvard University
2000-current. Standing Committee on Mind/Brain/Behavior, Harvard University
2002-2003, Subcommittee on Undergraduate Curriculum, Harvard Mind/Brain/Behavior Interfaculty
Initiative
2002. NIH National Institute on Deafness & Other Communication Disorders. Special Emphasis Review
Panel, Exploratory/Developmental Research Program
2003-4,2008-9, NSF Grant Review Committee
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Mark Judc Tramo. April 2009
2004-2007, Large Animal Users Group. Massachusetts General Hospital
2006-2009, Steering Committee, Mind Brain & Behavior Interfaculty Initiative, Harvard University
Professional Societies
I 983-current. American Academy of Neurology. Member
1985-current, American Society for Composers, Authors, and Publishers. Writer Member
I 986-current, Acoustical Society of America, Member
1989-current, Society for Neuroscience, Member
1991-1992, Stroke Council, American Heart Association, Fellow
1992-2007, American Association for the Advancement of Science
1995-current, Society for Music Perception & Cognition
1998-2008 Association for Research in Otolaryngology
2000-2008. New York Academy of Sciences
2006-7, Boston Music Research Association
2009-current, National Academy of Recording Arts & Sciences
Editorial Boards
1989-current, Reviewer, Science. Nature Neuroscience Reviews. Neuron, Journal of Neurophysiology. Trends
in Neuroscience& Journal of the Acoustical Society of America, Cerebral Cortex, Oxford Press (partial list)
1994-1998, Consulting Editor. Music Perception
1997-2005, Associate Editor, Journal of Cognitive Neuroscience
2007-current, Advisory Board. Journal of Interdisciplinary Musk Studies
Awards and Honors
1974-1978. Joseph Tauber Scholar. Yale University
1978. Bachelor of Arts cum laude with Honors in the Major (Biology), Yale University
1978-1982, Joseph Collins Foundation Scholar, Cornell University Medical College
1981-1982. Vincent Astor Scholar, Cornell University Medical College
1987-1988, National Research Service Award, Physiology, National Institute of Neurological &
Communicative Disorders & Stroke, National Institutes of Health
1988-1991. Program Project Award, Co-Principal Investigator, National Institute of Neurological Disorders &
Stroke. National Institutes of Health
1989-1991, Cognitive Science Award, Co-Principal Investigator, Office of Naval Research
1991-1992, National Research Service Award, National Institute of Mental Health
1992-1997, Clinical Scientist Development Award. National Institute on Deafness and Other Communication
Disorders, National Institutes of Health
1992-1993, Sackler Scholar in Psychobiology, Harvard University
1993-1994, Alwin M. Pappenheimer Scholar. Harvard University
1997, Research Award. National Organization for Hearing Research
1997-1998. Provost's Award for Educational Innovation. Harvard University
1997-2002, Cognitive Neuroscience Program Award, McDonnell-Pew Foundation
1998-2002. FIRST Award. National Institute on Deafness and Other Communication Disorders, National
Institutes of Health
2000-2001, Gordon and Llura Gund Research Award, Mind/Brain/Behavior Interfaculty Initiative, Harvard
University
2002-current, Institute for Music & Brain Science
2009, Performing Arts Medicine Association Richard Lederman Award
2009-2010, Grammy Foundation Research Award
PART II: Research, Teaching, and Clinical Contributions
A. Narrative Report of Research, Teaching, and Clinical Contributions
Research contributions include:
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• Experiments on auditory perception by stroke patients that corrected a 25-year old misconception, set
forth by Neff and colleagues in the 1975 Handbook of Sensory Physiology, and hitherto widely held by
auditory physiologists, psychologists, and audiologists, that auditory cortex is neither necessary nor
sufficient for simple pitch discrimination and loudness discrimination (Tramo 1990; Tramo et al. 2002:
Dykstra et al. 2008).
• Experiments on auditory perception by split-brain and stroke patients and event-related potential
experiments on auditory perception by normal adults that elucidated (a) the functional role of auditory
cortex in complex-pitch, harmony, and timbre perception: (b) the neuroanatomical dissociability of conical
mechanisms mediating the perception of harmony in its vertical vs. horizontal dimensions in the presence
vs. absence of tonal context; and (c) the location of interhemispheric auditory projections in the human
corpus callosum (Tramo et al. 1987. I 990. I991,1993,2005.2008ab; Tramo & Bharucha 1991; Tramo
& Koelsch 2008);
• Experiments on the neural coding of tonal harmony via intra-axonal and extracellular microelectrode
recordings of cat auditory nerve and inferior colliculus responses to complex-tones that: (a) elucidated
peripheral and central representations of tonal consonance and dissonance; and (b) provided a biologically-
based alternative to Helmholtz's counterintuitive theory, shared by many auditory physiologists and
psychologists for more than a century, that the consonance of isolated musical intervals and chords is a
negative percept. derived from the absence of tonal roughness; our neural data provide empirical evidence
that consonance is a positive percept. built up from multiple. harmonically-related pitches, in keeping with
the music-theoretic formulations of Rameau and others (Tramo et al. 1992.2001.2008a; McKinney et al.
2001);
• Experiments on auditory perception by neurological patients that elucidated the functional role of the
auditory midbrain (inferior colliculus) in spatial localization, speech recognition. and music recognition
(Litovsky et al. 2002; Tramo et al. 2008c);
• Experiments on the receptive field properties of macaque auditory cortex neurons via a novel alert
preparation for microelectrode recordings that avoided the confounding effects of anesthesia and,
consequently, uncovered a temporal response characteristics and spectrotemporal receptive field properties
(Tramo 1998; Tramo et al. I999a,b);
• Experiments on the neural coding of species-specific communication sounds in the primary auditory
cortex of alert rhesus monkeys that revealed a candidate coding scheme based on the coarse temporal
pattern of responses to different vocalization categories (Tramo 1998; Tramo et al. 1996. 1998);
• Experiments on auditory acuity and experience-dependent plasticity in musicians and non-musicians
that supports the claim that high perceptual acuity for music-related auditory functions is universal, and that
the "use it or lose it" principle applies to the effects of music experience (formal and informal) on high
acuity (Tramo et al. 2008d):
• Experiments with premature infants supporting the claims that (a) music-related auditory functions are
universal; and (b) controlled auditory stimulation with music ameliorates pain and stress caused by medical
procedures (Tramo a al. 2006; Lense et al. 2008);
• Development of novel MRI-based morphometric methods for flat-mapping and measuring the surface
of human cerebral cortex; application of the methods revealed prenatal influences on the morphology of
gyral-sulcal patterns in monozygotic twins and pathoanatomical correlates in patients with amusia,
dyslexia, and schizophrenia (Tramo et al. 1990, 1995, 1998; Jouandet et al. 1989; Green a al. 1991, 1999;
Loftus et al. 1995).
Teaching contributions include:
• Founding the Auditory• Neurobiology lecture course (Neurobiology 101) at Harvard College and
Graduate School of Arts & Sciences (2007-current);
• Founding the Music. Mind & Brain seminar course (Psychology 987b. then MBB 98) at Harvard
College as a Faculty Fellow of the Harvard University Mind/Brain/Behavior Interfaculty Initiative; this is
the longest running course in the history of the MBB Initiative (1995-current):
• Sponsoring the Honors Theses and Research Awards of nine Harvard College students (1995-current);
• Sponsoring Research Assistantships for over a dozen Harvard College, HMS, MIT. and outside students
(1994-current);
• Serving on thesis advisory and qualifying examination committees and sponsoring research and student
laboratory rotations as a faculty member of the HST Speech & Hearing Bioscience & Techology Graduate
Studies Program
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Mark Judc Tram°. April 2009
• Co-Founding, Directing, and lecturing in the annual Harvard Medical School Continuing Medical
Education course. Dementia: A Comprehensive Update (1997-current); during my turn as Director in 2001.
I revised the course in collaboration with the Co-Directors and doubled its registration, thereby reversing a
downward trend that caused some to consider discontinuing the course or changing the topic: since then,
we have consistently received the Dean of the HMS Dept of CME's commendation as one of their highest
rated courses; during my 2007 turn as Director we set a new record for registration.
• Co-founding the Music Perception and Cognition seminar course (HST 725) in the Harvard-M.I.T.
Speech & Hearing Biosciences & Technology Graduate Studies Program (2004.2(x7)
• Supervising laboratory research and independent studies for over two dozen undergraduate students.
graduate students, medical students, and neurology residents from Harvard, M.I.T., and outside institutions
(1994-current);
• Organizing symposia, serving on fellowship and grant application committees, and helping to build the
MBB curriculum at FAS and GSAS as a member of the Harvard University Steering Committee on
Mind/Brain/Behavior (2006-current) and the Standing Committee on Neuroscience/MBB (2000-current);
• Organizing the Cognitive/Behavioral Neurology teaching curriculum at Massachusetts General Hospital
for second-year Partners Neurology Residents (1995-2006):
• Professional education via invited CME lectures and Neurology Grand Rounds (1998-current)
• Public education via my contributions to and recorded appearances in the science museum exhibit, Wild
Music!, which is touring the United States through at least 2009. and through public lectures (e.g., Carnegie
Hall, Lincoln Center), materials posted on The Institute for Music & Brain Science website, and interviews
published by print, radio, television, and Internet journalists around the world;
• Middle- and high-school education via my contributions to the interactive edutainment DVD,
Neuromairix. winner of the Robert Wood Johnson Foundation's 2007 Games for Health Competition.
Clinical contributions include:
• One General Neurology Inpatient Service rotation per year at Massachusetts General Hospital (1995-
current);
• Outpatient Service in the Cognitive/Behavioral Neurology Unit at Massachusetts General Hospital
(1992-current);
• Inpatient and Outpatient Behavioral Neurology Consults at McLean Hospital (1996-2005).
B. Funding Information
Research Funding
1987-1988, National Institute of Neurological & Communicative Disorders & Stroke. National Institutes of
Health. Fellow. National Research Service Award (Physiology), "Metabolic Correlates of the
Neglect Syndrome". NS08159
1988-1991, National Institute of Neurological Disorders & Stroke. National Institutes of Health. Co-PI.
Program Project in Cognitive Neuroscience, Project III: "Cortical Networks in Auditory Pattern
Perception" and Project VIII: "Core Neurology and Brain Imaging". NS 17778
1989-1991, Office of Naval Research. Cognitive Science Program Award , "Brainprints: Straight-line two-
dimensional maps of the human cerebral cortex in vivo', N00014-894-3035
1991-1992, National Institute of Mental Health. Fellow, National Research Service Award. Program in
Neuroscience, MH18012
1992-1997, National Institute on Deafness and Other Communication Disorders. National Institutes of
Health, PI, Clinical Scientist Development Award. "Functional Anatomy of Auditory Cortex".
DC00071
1993-1995. National Institute of Mental Health. Consultant. Collaborative Biological Research in
Schizophrenia, MH31154
1997-2002, McDonnell-Pew Foundation, PI. Cognitive Neuroscience Program Award, "Neural Coding of
Communication Signals in Auditory Cortex", 97-25
1997-1998, National Organization for Hearing Research, P1. Research Award, 'Physiology of Neuronal
Responses to Vocal Communication Sounds in the Auditory Cortex of Awake-Behaving Macaques".
1998.2002, National Institute on Deafness and Other Communication Disorders, National Institutes of
Health. PI. First Independent Research Support and Transition Award. "Physiology of Vocalization
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Sensitivity in Auditory Cortex". DC03382
2000-2001, Gordon and Llura Gund Research Award, Mind/Brain/Behavior Initiative, Harvard University,
PI, "How the brain evolved speech representations: Field experiments, neurophysiology, and fMRI".
2001-2006. Peabody Charitable Foundation. Co-Investigator, "Auditory Neurology"
2002-current, The Institute for Music & Brain Science. Principal Investigator, "Program in Auditory
Neuroscience"
2009-current, Grammy Foundation Research Award, Principal Investigator, "Treatment of latrogenic Pain and
Stress in Critically-Ill Premature Infants"
Education Funding
2004-2006. National Institute for Mental Health Small Business Innovative Research Award. Morphonix,
Inc., Consultant, "Neuromatrix: Interactive Neuroscience Education", MH066429
2005-2009, National Science Foundation Informal Science Education Award, Consultant, "Wild Music!"
Museum Exhibit and Tour, ESI-0211611
2007-2008, National Endowment for the Humanities, Digital Humanities Grant, Consultant."uTunes: Music
1.01". HD-50258-07
C. Report of Current Research Activities
I am currently active in data collection, data analysis, and/or manuscript preparation of original research
with colleagues and students on: I) effects of unilateral and bilateral auditory cortex lesions on virtual pitch
perception and inharmonicity detection by stroke patients; 2) localization and latency of cortical auditory
event-related potentials evoked by harmonic context violations during consonance judgments; 3) fMRI
activation of auditory cortex subregions during aesthetic evaluation of harmonic progressions; 4) effects of
film music on affective evaluation of facial expressions; 5) effects of controlled auditory stimulation with
music, speech, and natural sounds on physiological and behavioral indices pain and stress in premature
infants.
D. Report of Teaching
I. Local contributions
a. Medical SchooVDental School/Graduate School
Cornell University Medical College
1987-1988, Third Year Clerkship in Neurology, Tutor, 4 students/yr, 20 hrs/yr
1987-1988. Fourth Year Neurology Elective in Cognitive Neuroscience. Tutor, 2 students/yr, 40 hrs/yr
1987-1988, First Year Student Faculty Advisor, 6 students/yr, 24 hrs/yr
Dartmouth Medical School
1989-1991, McDonnell-Pew Graduate Studies in Cognitive Neuroscience. Lecturer & Laboratory
Instructor, 2 students/yr, 60 hrs/yr
1998-1991, Scientific Basis of Medicine: Neurology, Lecturer, entire class of 2nd yr students/yr, 10
hrs/yr
1989-1991, Fourth Year Elective in Cognitive Neuroscience. Tutor. 2 students/yr. 40 hrs/yr
Harvard Medical School. Division of Medical Sciences, and Division of Health Sciences and Technology
1993. Second Year Physical Diagnosis in Neurology, Tutor, 4 students/yr, 16 hrs/yr
1993. HST 722: Brain Mechanisms of Hearing and Speech, Lecturer. 12 graduate students/yr. 4 hrs/yr
1993. 1995. 1997, Neurobiology 208: The Visual System, Lecturer, 10 medical and graduate
students/yr, 6 hrs/yr
1994-current. Clinical Clerkship in Neurology. Examiner, I medical student/yr. 2 hrs/yr
1996.1999,2007, HMS/HDS Human Nervous System & Behavior. Lecturer & Tutor, 12.20
medical/dental students/yr 60 hrs/yr
1997,1998,2005 HST 799, Special Topics Research in the Speech & Hearing Sciences, 1 student/yr,
120 hrs/yr
1999-2000, Thesis Advisory Committee, HST Speech & Hearing Sciences, I graduate student/yr, 16
hrs/yr
2000. Thesis Qualifying Examination Committee. HST. Speech & Hearing Sciences, I graduate
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student/yr, 4 hrs
2004. 2007. HST 725. Music Perception & Cognition (Co-Instructor). 4-8 students/semester
b. Harvard College. Harvard Graduate School of Arts & Sciences, and the Harvard University
Mind/Brain/Behavior Interfaculty Initiative
1995-2007. Psychology 985. Preparation for the Honors Thesis, I undergraduate every other yr, 50 hrs/student
1995-current, Psychology 990, Honors Thesis Advisor, I undergraduate every other year, 100 hrs/student
1997-2000. Psychology 910r. Supervised Reading/Research, I undergraduate every other year, 50 hrs/student
1998, 2002 Biology 98r, Introduction to Research. I undergraduate/yr, 50 hrs/yr
1998-1999, Biology 99, Honors Thesis Advisor, 2 undergraduates/yr, 100 hrs/student
1998, 2000. 2001. Psychology 987b. Music. Mind. & Brain, 12 undergraduates/yr. 100 hrs/yr
1999-2001, Mind/Brain/Behavior Thesis Workshop. 12 undergraduates/yr. IS hrs/yr
2000-20001, Psychology Honors Thesis Committee, 2 undergraduates/yr; 20 hrs/student
2000-2001, Special Concentrations 99, Honors Thesis Advisor, I undergraduate/yr, 100 hrs/yr
2001-2002, Special Concentrations 91r, Supervised Reading and Research, I undergraduate, IS hrs/yr
2002-2003. PhD Thesis Committee, Cory Miller, Psychology. 20 hrs/yr
c. Graduate Medical Courses
The New York Hospital - Cornell Medical Center
1986-1988, Neurology Outpatient Clinic, Attending, PGY 2.4 Neurology Residents, 48 hrs/yr
1986-1988, Cognitive Neuroscience Seminar Series for Neurology Residents. Organizer and Lecturer.
36 hrs/yr
1987-1988. Neurology Consult Service. Attending. PGY-3 Neurology Resident. 112 hrs/yr
Dartmouth-Hitchcock Medical Center
1988-1991, Neurology Outpatient Clinic, Attending, PGY 2-4 Neurology Residents. 60 hrs/yr
1988-1991. Resident Research Elective in Cognitive Neuroscience, Tutor. 40 hrs/yr
1989.1991, Behavioral Neurology Ward Rounds, 12 hrs/yr
Massachusetts General Hospital
1992-current, Neurology Ward or Consult Service, Attending, PGY I.2, and 4 Neurology,
Psychiatry, and Physiatry Residents and Interns, 224 hrs/yr
1992-current, Cognitive/Behavioral Neurology Unit. Attending, PGY 3 Neurology Residents,
48 hrs/yr
1992-current. Neurology Residents Cognitive/Behavioral Neurology Seminar. Attending. 24 hrs/yr
1994. 2000. Neurology Residents Neuroscience Seminar. Lecturer, PGY 2-4 Neurology Residents, 5
hrs/yr
1995-current, General Neurology Residents' Clinic, Attending. PGY 2-4 Neurology Residents.
30 hrs/yr
1998. 1999. 2001 . Neurology Professor's Rounds. PGY 1-4 Neurology, Psychiatry, and Physiatry
Residents and Interns, 2 hrs/yr
1999, 2000. 2005, 2006 Behavioral Neurology Signs Rounds. PGY 1-4 Neurology, Psychiatry, and
Physiatry 12 hrs/yr
Residents and Interns, 2-4 hrs/yr
2000-current. Mortality and Morbidity Conference, PGY 2-4 Neurology Residents, 18 hrs/yr
d. Invited Presentations
The New York Hospital - Cornell Medical Center:
1986-1988, Neurology/Neurosurgery Grand Rounds, Lecturer, 20 hrs/yr
Dartmouth-Hitchcock Medical Center:
1989.1991, Neurology/Neurosurgery Grand Rounds, Lecturer, 20 hrs/yr
Harvard Medical School
1992. Program in Neuroscience Seminar, Wood's Hole, Lecturer, 20 hrs
1993, Neurobiology Department Seminar Series, Lecturer, 20 hrs
2006. Medical Exchange Club, Lecturer, Harvard Club of Boston. 2 hrs
Harvard University
1994, Multiple Perspectives in Music Seminar Series. Music Department, Lecturer, 10 hrs
1998. Psychology Department Seminar Series. Lecturer, 20 hrs
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2000. Mind/Brain/Behavior Faculty Fellowship Seminar Series, 20 hrs
2001. Harvard University Mind/Brain/Behavior Advisors' Dinner, Lecturer, 21 Club, New York, 5 hrs
2001, The Harvard Club of Boston, A Saturday of Symposia, Music. Mind, and Brain (invited)
2001. Mind/Brain/Behavior Advisory Board Seminar. 20 hrs
Massachusetts General Hospital
1997,2001. Neurology Grand Rounds. Lecturer. 10 hrs/yr
Brigham & Women's Hospital
1994, Behavioral Neuroscience Seminar Series, Lecturer, 20 hrs
Beth Israel Hospital
1993, Neurology Grand Rounds, Lecturer, 1993,20 hrs
Cambridge Hospital
2000,2001. Neurology Lecture Series. 10 hrs/yr
e. Continuing Medical Education Courses
Cornell Medical College
1988. Review of Internal Medicine. Lecturer. 20 hrs
Harvard Medical School
1997-2000,2002-2006, Cognitive and Behavioral Neurology: Focus on Dementia. Co-Director and
Lecturer. 40 hrs/yr
2001,2007 Dementia: A Comprehensive Update, Director and Lecturer, 100 hrs
f. Advisory and Supervisory Responsibilities
Cornell Medical College
1987-1988, First Year Student Faculty Advisor. 6 students/yr, 24 hrs/yr
Massachusetts General Hospital
2006-2007. Mentor, Senior Neurology Resident (I), 50 hrs/yr
2004-8-current, Preceptor. Neurology Residents Clinic (200 hrs/yr)
2. Regional, national, or international contributions
a. Presentations
1985, Session on Multiple Sclerosis: Clinical Investigation and Therapy, 37th Annual Meeting of the
American Academy of Neurology
1987. Cerebrovascular Disease II: Poster Presentations, American Neurological Association 114th Annual
Meeting
1988. Session on Cerebral Metabolism. 40th Annual Meeting of the American Academy of Neurology
1989, Behavioral Neurology Poster Session, American Neurological Association 114th Annual Meeting
1989. Interhemispheric Relations Poster Session, Society for Neuroscience 19th Annual Meeting
1989. McDonnell Foundation Summer Institute in Cognitive Neuroscience Faculty and Lecturer (invited)
1990, Second International Conference on Music and the Cognitive Sciences, Cambridge University. Lecturer (invited)
1990. Human Behavioral Neurobiology Poster Session, Society for Neuroscience 20th Annual Meeting
1990, First Annual McDonnell-Pew Program in Cognitive Neuroscience Investigators' Workshop, Lecturer (invited)
1990, Behavioral Neurology Poster Session, American Neurological Association 115th Annual Meeting
1990, McDonnell Foundation Summer Institute in Cognitive Neuroscience Faculty and Lecturer (invited)
1991, Functional Aspects of Auditory Cortex Symposium, Lecturer, International Brain Research Organization
Third World Congress of Neuroscience, (invited)
1991, Music and the Brain Session, Lecturer, Second International Conference on Music Perception and Cognition,
UCLA (invited)
1991, Central Auditory Physiology Poster Session, Society for Neuroscience 21st Annual Meeting
1991. Cognitive Neuroscience Session. Co-Chair and Lecturer. 2nd International Conference on Epilepsy and the
Corpus Callosum (invited)
1991, Scientific Session, Lecturer, Society for University Neurosurgeon (invited)
1991. Neurology Grand Rounds. Lecturer, Yale School of Medicine Neurological Study Unit and Yale-New Haven
Hospital (invited)
1992, Central Auditory Physiology Slide Session. Society for Neuroscience 22nd Annual Meeting
1992. Music and the Brain Symposium. Lecturer. Art Institute of Chicago (invited)
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1992. International Conference on the Psychophysiology and Psychopathology of the Sense of Music. Lecturer (invited)
1993, Central Auditory Physiology Slide Session, Society for Neuroscience 23rd Annual Meeting
1994, Central Auditory Physiology Poster Session, Society for Neuroscience 24th Annual Meeting
1996. Functional Aspects of Auditory Cortex Workshop, Lecturer. XXVIth International Congress of Psychology
(invited)
1996. Auditory Cortex Physiology - Primates Poster Session, Society for Neuroscience 26th Annual Meeting
1997, Music and the Brain Symposium, Lecturer, International Computer Music Conference, (invited)
1997, Pedagogical Session: Neurobiology , Lecturer, International Conference on Complex Systems
(invited)
1998, Press Conference on Music and the Brain, Moderator, Society for Neuroscience 28th Annual
Meeting (invited)
1998, Auditory Cortex Physiology - Complex Sounds Poster Session, Society for Neuroscience 28th
Annual Meeting
1999, Auditory Cortex Slide Session, 22nd Association for Research in Otolaryngology Midwinter Meeting
1999, Central Auditory Physiology: Spectrotemporal Coding Poster Session, Society for Neuroscience
29th Annual Meeting
2000, Biological Foundations of Music. Lecturer, New York Academy of Sciences (invited)
2000. The Music of Nature and the Nature of Music. Lecturer. Smithsonian Institute, National Musical
Arts. National Academy of Sciences (invited)
2000. Association for Research in Otolaryngology: Inner Ear: Normal Structure or Function Poster Session
2001. Auditory: Coding. Tuning and Perception Poster Session. Society for Neuroscience 31st Annual
Meeting
2001, Bcrklee College of Music. Music Therapy Symposium, Music & The Brain (invited)
2002, Learning & The Brain Symposium, Music Education, & The Brain. Cambridge, MA (invited)
2002, Hudson Theatre, Beth Abraham Music Therapy Program, Music Cognition. NY, NY (invited)
2002. Auditory Neuroscience Seminar Series, University of Connecticut at Storrs, Effects of Auditory
Cortex Lesions on Music Perception (invited)
2003, Neuroesthetics Symposium, University of California at Berkeley, Widely Distributed Neural Systems
for Musical Aesthetics (invited)
2004, Linda and Jack Gill Center for Biomolecular 2004 Symposium and Award, Indiana University (invited)
2004, Jill S. Titus Lecture. Institute for Mind. Body & Spirituality, Lesley College (invited)
2004, Special Session of the 50' Annual Meeting of the Acoustic Society of America, Musical Acoustics:
Neurophysiology of Musical Instrument Playing. New York (invited)
2004, Carnegie Hall Concert and Symposium, Beautiful Minds, Beautiful Music, New York (invited)
2005. Neurosciences & Music Symposium. Leipzig, Germay (invited)
2005. Leibniz Institute. Magdeburg Germany (invited)
2005, Harvard Club of Fort Lee. FL (invited)
2005. Roland M. Pinkham Annual Memorial Lecture, Swedish Medical Center, Seattle. WA (invited)
2006. Neurosciences Institute. LaJolla. CA (invited)
2006, Arts in Medicine Series. Memorial Sloan-Kettering Cancer Center (invited)
2007. National Center for Human Performance. Houston. TX (invited)
2007, Center for Performing Arts Medicine, Methodist-Cornell Medical Center, Houston, TX (invited)
2007, Cold Spring Harbor InterdisciplinarySymposium on Memory in Neuroscience & the Humanities
(invited)
2008, National Academy of Sciences Annual Meeting. Koshland Science Museum, Music and the Brain:
Performance and Discussion. Washington. DC (invited)
2009, Cleveland Clinic, Keynote Lecture. Music ad the Brain Symposium, Cleveland, OH (invited)
2008, Duke University, Opening Lecture. Music and the Brain Sympsosium, Durham, NC (invited)
2009, Shepherd School of Music, Rice University, Exploring the Mind Through Music Symposium,
Houston. TX (invited)
2009. Barrow Neurological Institute. Neurology/Neurosurgery Grand Rounds. Phoenix. AZ (invited)
2009. International Mind Brain & Education Society Meeting. Panel: Advances in the cognitive
neuroscience of music: Implications for education. Philadelphia. PN
2009. International Mind Brain & Education Society Meeting. Panel: Emotion, learning. & the brain:
Insights from experimental neuroscience & educational approaches to children with neurological
disease, Philadelphia. PN
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2009. Brigham & Women's Hospital and Massachusetts General Hospital. Michael Ty Memorial Lecture,
Boston, Massachusetts
2009, Performing Arts Medicine Association, Richard J. Lederman Keynote Lecture. Snowmass, CO
2009. Lincoln Center, Music and the Brain Symposium, Closing Lecture, New York. NY
3. Teaching Awards
1997-1998. Harvard University Provost's Award for Educational Innovation
2000. Harvard Medical School Class of 2000 Faculty Award for Excellence in Clinical Teaching
4. Description of major curriculum offerings and innovative programs
Founded interdisciplinary Harvard College course, "Music Mind & Brain" (Psychology 987b:
Mind/Brain/Behavior Interfaculty Initiative) in 1995. Eleventh offering this coming Spring Term of
2007-2008 Academic Year.
Proposal for new undergraduate Auditory Neurobiology course recommended for approval by the FAS
Standing Committee on Neurobiology in the Fall Term of the 2006-2007 Academic Year: final
approval by FAS pending for initial offering in the Fall Term of the 2007-2008 Academic Year as
a Neurobiology course administered by the Department of Molecular and Cellular Biology.
Sponsored seven Harvard undergraduate honors theses in Biology, Psychology, and Special Concentrations
from 1996-2006, with one additional student actively working on her 2008 thesis in Human
Evolutionary Biology.
Founded the world's first non-profit organization, The Institute for Music & Brain Science, devoted to
basic and clinical research on music, science, and medicine, in collaboration with four Harvard
Medical School colleagues, The Beatles' producer, Sir George Martin, and several colleagues from
academia, the arts, and industry.
Appointed principal tutor of MGH Cognitive Neurology Unit in 1995, revised teaching curriculum
Recipient of the Harvard Provost's Award for Educational Innovation, 1997-1998 Academic Year.
5. Public Education (partial list)
1995. NBC Nightly News with Tom Brokaw, The New York Times Science Times, Boston Magazine
1996, BBC 4 Television, Los Angeles Times. Atlanta Journal-Constitution, Young Students Learning Library
Science Yearbook, American Psychological Association Monitor
1997, The Washington Post, The Boston Globe Quotes of Note, Fox News Channel
1998, NBC Today Show, PBS Healthweek, CBS National Radio, Public Radio International. New England
Cable News, KCBS Radio San Francisco. WAV-FM Radio Tokyo. ZIP-FM Radio Nagoya. LA Times.
USA Today
1999. Lifetime Channel, Discovery Science Channel, NPR Living On Earth
2001, National Geographic TV, WBZ-TV Boston, WCVB-TV Boston, Discovery Channel, NPR The
Connection, US News & World Report, Discover Magazine, The New York Times Science Times,
The Boston Globe Health/Science
2002, WCVB-TV Boston, Canadian Broadcasting Corporation Radio Ideas, People Magazine. ABCnews.com,
NPR Infinite Mind
2003, The New York Times. Harvard Medical School Alumni Bulletin, Neurology Today. Edmonton Journal
2004, Journal of the American Medical Association, PBS-TV Closer To Math, WiredNews.com, Discover
2005, CBS-TV Sunday Morning. National Public Radio, Chicago Tribune, Seattle Times
2006, The Wall Street Journal, US News & World Report, NPR, St Louis Post-Dispatch
2007. Washington Post
2008. Nightline, WABC-TV
2009, All Things Considered, NPR
E. Report of Clinical Activities
1. Description of Clinical Practice
I am a general adult neurologist (board certified in 1988) with fellowship training and 20 years of outpatient
and teaching experience in the subspecialty of Behavioral Neurology, which deals with disorders of memory.
language, attention, auditory cognition, visual cognition, perceptual-motor integration, reasoning, learning. and
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emotion caused by stroke, neurodegenerative disease, head trauma, systemic diseases, and developmental
diseases.
2. Patient Load and Complexity
I see up to eight patients every other week in the MGH Cognitive Neurology Outpatient Unit. I typically serve
as the Attending on one of the Neurology Inpatient Services one month per year, with 10 days per month of
Emergency Room coverage during that month.
PART Ill.
A. Bibliography
I. Tramo MJ, Hainline B, Petito F, Lee B, Caronna .1 Vertebral artery injury and cerebellar stroke
while swimming. Stroke 1985; 16:1039-1042
2. Tramo MJ, Schneck MJ, Lee BCP, Rapoport S Evoked potentials and magnetic resonance imaging in
3. Tramo MJ, Hainline B Stroke in sports. In: Jordan B, Tsairis P, Warren R, eds. Sports Neurology.
Rockville, Maryland: Aspen Publishing Co. 1989:98-115
the diagnosis of multiple sclerosis. Neurology 1985; 35(Suppl I):105
4. Levy DE, Sidtis J, Rottenberg DA, Jarden JO, Strothers S, Dhawan V, Ginos JZ, Tramo MJ, Evans
AC, Plum F The vegetative state and locked-in states affect cerebral blood flow and glucose
utilization differently. Annals of Neurology 1987; 22:673.682
5. Tramo MJ, Sidtis LI, Dhawan V, Strothers S. Moeller !, Ginos J, Sergi M, and Rottenberg DA
Variability of regional glucose extraction fraction in the normal resting state. Annals of
Neurology 1987; 22(Suppl):16 1 A
6. Tramo MJ, Gazzaniga MS Discrimination and recognition of complex tonal spectra by the cerebral
hemispheres: Differential lateralization of acoustic-discriminative and semantic-associative
functions in auditory pattern perception. Society for Neuroscience Abstracts 1987; 15:1060
7. Tramo MJ, Baynes K, Volpe B Impaired syntactic comprehension and production in Broca's aphasia:
CT lesion localization and recovery patterns. Neurology 1988; 38:95-98
8. Eidelberg D, Tramo M, Strother SC, Moeller JR, Sidtis iJ, Dhawan V, Rottenberg DA Variability in
regional cerebral metabolic rate for glucose (rCMRGIu) and the consequences for the
study of neurologic disease with FDG/PET. Neurology 1988; 38(Suppl I ):367
9. Tramo MJ, Gazzaniga MS Recovery patterns in neurobehavioral syndromes. W.H.O. Collaborating
Centre for Research and Training in Neurosciences, Neuroplasticity of the Nervous System, Beijing,
10. Jouandet ML, Tramo MJ, Herron DM, Hermann A, Loftus WC, Bazell J, Gazzaniga MS Brainprints:
Computer-generated two-dimensional maps of the human cerebral cortex in vivo. Journal of
Cognitive Neuroscience 1989; 1: 88-117
II. Oppenheim IS. Skerry JE, Tramo MI, Gazzaniga MS Magnetic resonance imaging morphology of the
corpus callosum in monozygotic twins. Annals of Neurology 1989; 26:100-104
12. Levy DE, Tramo MJ, Plum F Do positron emission tomographic scans measure quality of life?
Reply. Annals of Neurology 1989; 2:288
13. Tramo MI, Reuter-Lorenz P, Gazzaniga MS Pure alexia: Cognitive and anatomical correlates. Annals
of Neurology 1989; 26:126
14. Tramo MI. Bharucha JJ, Musick FE Music perception and cognition following bilateral lesions of
auditory cortex. Journal of Cognitive Neuroscience 1990; 2:195-212
(Abstracted by permission In: Hall III, J.. ed. /991 Year Book of 'Speech. Language, and Hearing,
Chicago: Mosby 1991; Chapter 15: Auditory Findings in Pathologies)
15. Jouandet ML, Tramo MJ, Thomas CE, Newton CH, Loftus WC, Weaver JB, Gazzaniga MS
Brainprints: Inter- and intra-observer reliability. Society for Neuroscience Abstracts 1990; 16:1151
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16. Tramo MI. Guglielmo MA, Reuter-Lorenz P. Gazzaniga MS Functional dissociations in
hemispatial neglect: Brainprints and quantitative lesion localization. Annals of Neurology 1990;
28:255
17. Tramo MI Current Problems in Neurology: 5, Impact of Functional Imaging in Neurology and
Psychiatry: Book Review. Neurosurgery 1990; 26:354-355
IS. Tramo MJ Lesion Analysis in Neuropsychology: Book Review. Journal of Cognitive Neuroscience
1990;2:156-157
19. Tramo MJ, Bharucha JJ Musical priming by the right hemisphere post-callosotomy.
Neuropsychologia 1991; 29:313-325
20. Tramo MJ, Bharucha JJ Conical networks in associative auditory processing. International Brain
Research Organization Abstracts 1991; 252
21. Tramo MI, Musiek FE, Gazzaniga MS Disruption of interhemispheric integration of complex auditory
information following focal hemorrhage into the posterior body of the corpus callosum. Society for
Neuroscience Abstracts 1991; 17:1484
22. Zatorre RJ, Heffner H, Liegeois-Chavel C, Tramo M, Samson S Functional aspects of auditor),
cortex. International Brain Research Organization Abstracts 1991; 8
23. Green RL, Tramo MJ, Loftus WC, Thomas CE, Brown PB, Weaver JB, Gazzaniga MS
Regional cortical surface area measurements in monozygotic twins discordant for schizophrenia
suggest a left hemisphere basis for the disease. Society for Neuroscience Abstracts 1991; 17:455
24. Baynes K, Tramo M.L. Gazzaniga MS Reading with a limited lexicon in the right hemisphere of a
callosotomy patient. Neuropsychologia 1992; 30:187-200
25. Kussmaul CL, Tramo MJ, Mangun GR Investigation of harmonic relationships on auditory event-
related potentials to successive pure tones. Psychophysiology 1992; 29:547
26. Baynes K, Tramo, MJ, Fendrich R, Reeves AG, Gazzaniga MS Specificity of interhemispheric
transfer following a partial lesion of the corpus callosum. Society for Neuroscience Abstracts 1992;
18:1207
27. Tramo MJ, Cariani PA, Delgutte B Representation of tonal consonance and dissonance in the
temporal firing patterns of auditory nerve fibers: Responses to musical intervals composed of pure
tones vs. harmonic complex tones. Society for Neuroscience Abstracts 1992; 18:382
28. Tramo M1 Split-brain studies of music perception and cognition. Contemporary Music Review 1993;
9:113-121
29. Loftus WC, Tramo MI, Thomas CE, Green RL, Nordgren RA. Gazzaniga MS Three dimensional
quantitative analysis of hemispheric asymmetry in the human superior temporal region. Cerebral
Cortex 1993:3:349-355
30. Delgutte B, Cariani PA, Tramo MJ Neurophysiological correlates of the pitch of complex tones.
Journal of the Acoustical Society of America, Suppl. 1993; I, 93:2293-2294
31. Bharucha ii, Tramo MJ, Zatorre RJ Abstraction of the missing fundamental following bilateral
lesions of auditory cortex. Society for Neuroscience Abstracts 1993; 19:1687
32. Peretz 1, Kolinsky R, Tramo M, Labrecque R, Hublet C, Demeurisse G, Belleville S Functional
dissociations following bilateral lesions of auditory cortex. Brain 1994; 117:1283.1301
33. Tramo MJ, Grant A, Braida LD Psychophysical measurements of frequency difference limens for
relative pitch discrimination reveal a deficit following bilateral lesions of auditory cortex. Society
for Neuroscience Abstracts 1994; 20: 325
34. Kussmaul CL, Tramo MJ, Mangun GR ERP measures of musical expectancy: Role of the NI 00 and
other early components. Proceedings of the Herbert von Karajan Symposium, Wien, 1994
35. Tramo MI, Loftus WC, Thomas CE. Green RL, Mott LA, Gazzaniga MS Surface area of human
cerebral cortex and its gross morphological subdivisions. In vivo measurements of monozygotic
twins suggest differential hemisphere effects of genetic factors. Journal of Cognitive Neuroscience
1995; 7:292-302
36. Loftus WC, Tramo MJ, Gazzaniga MS Conical surface modeling reveals gross morphometric
correlates of individual differences. Human Brain Mapping 1995; 3:257-270
37. Tramo MI, Baynes K, Fendrich R, Mangun GR, Phelps EA, Reuter-Lorenz. PA, Gazzaniga MS
Hemispheric specialization and interhemispheric integration: Insights from experiments with
commissurotomy patients. In: Reeves AG, Roberts DW, eds., Epilepsy and the Corpus Callosum II, pp
263-295, NY: Plenum Press, 1995
38. Tramo MJ, Bellew BF, Hauser MD Discharge patterns of auditory conical neurons evoked by species-
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specific vocalizations and complex synthetic signals in Macaca mtdatta. Society for Neuroscience
Abstracts 1996; 22:1623
39. Tramo MJ Neural representations of tonal harmony. International Journal of Psychology 1996;
31:187
40. Baynes K, Tramo MJ, Reeves AG, Gananiga MS Isolation of a right hemisphere cognitive system in
a patient with anarchic (alien) hand syndrome. Neuropsychologia 1997; 35:1159-1173
41. Cariani P, Tramo M. Delgutte B Neural representations of pitch through temporal autocorrelation.
Proceedings of the Audio Engineering Society, AES Preprint #4583 (L-3), 1997
42. Tramo MJ, Loftus WC, Stukel TA, Green RL, Weaver JB, Garaniga MS Brain size, head size, and
intelligence in monozygotic twins. Neurology 1998; 50:1246-1252
43. Patel AD, Peretz I, Tramo M, Labreque R Processing prosodic and musical patterns: A
neuropsychological investigation. Brain and Language 1998; 61:123-144
44. Tramo MJ, Moraru D, Hauser MD Representation of species-specific vocalizations in the temporal
discharge patterns of auditory conical neurons in alert Macaca mulatto. Society for Neuroscience
Abstracts 1998; 24:401
45. Tramo Mt Physiological acoustics. In: Beyer RT, Parker SP, eds.. McGraw-Hill 1998 Yearbook of
Science and Technology, Encyclopedia of Science and Technology, NY:McGraw-Hill , pp 305-307
46. Green RL. Hutsler ii, Loftus WC. Tramo MI, Thomas CE. Silberfarb AW, Nordgren RA, Gazzaniga
MS. The caudal infrasylvian surface in dyslexia. Neurology 1999; 53:974-981
47. Tramo MI, Cariani PA Receptive field organization in the core area of alert macaque auditory cortex.
Society for Neuroscience Abstracts 1999; 25:395
48. Tramo Mt. Cariani PA, Moraru D Spectrotemporal response properties of auditory cortex neurons in
the alert macaque. Association for Research in Otolaryngology Abstracts 1999; 22:116-117
49. Kass JH, Hackett TA, Tramo MJ Auditory processing in primate cerebral cortex. Current Opinion in
Neurobiology 1999; 9:164-170
50. Tramo MI, McKinney MC, Cariani PA, Delgutte B Physiology of tonal consonance and dissonance.
Association for Research in Otolaryngology Abstracts 2000; 23:275-276
51. Litovsky R, Fligor B, Tramo M Functional role of the human inferior colliculus in binaural hearing.
Association for Research in Otolaryngology Abstracts 2000; 23: 286
52. Tramo MI Music of the hemispheres. Science 2001; 291:54-5625.
53. Tramo MJ, Cariani PA, Delgutte B, Braida LD Neurobiological foundations for the theory of harmony
in Western tonal music. Annals of the New York Academy of Sciences 2001; 930:92-116
54. Battelli L, Cavanagh P. Intriligator J, Tramo MJ, Henaff M-A, Michel F, Barton DS Unilateral right
parietal damage leads to bilateral deficit for high-level motion. Neuron 2001; 32:985-995
55. McKinney MF, Tramo, MJ, Delgutte B Neural correlates of the dissonance of musical intervals in the
inferior colliculus. Association for Research in Otolaryngology Abstracts 2001; 24:54-55
56. Tramo MJ, Rosenbaum ER, Cariani PA, Hauser MD Differential responses of auditory cortex neurons
to natural and unnatural vocalizations in alert Macaca mulatto.
Society for Neuroscience
Abstracts 2001; 26
57. McKinney MF, Tramo, MJ, Delgutte B
Neural correlates of musical dissonance in the inferior
colliculus. In Houtsma AIM, Kohlrausch A, Prijs VF, Schoonhoven R, Eds., Physiological and
Psychophysical Bases of Auditory Function, Maastricth: Shaker Publishing, pp 71-77, 2001
58. Gray PM, Payne R, Krause B, Tramo MJ The biology of music. Response. Science 2001; 292:2432-
2433
59. Tramo MJ Pathoanatomy of amusia: Harmony perception and aesthetics. Journal Watch Neurology
2001; 3:88
60. Tramo MJ, Shah GD, Braids LB Functional role of auditory cortex in frequency processing
and pitch perception. Journal of Neurophysiology 2002; 87:122-139
61. Litovsky RY, Fligor B, Tramo MI Functional role of the inferior colliculus in binaural hearing.
Hearing Research 2002; 165; 177-188
62. Tramo MJ The Brain and Music. Harvard Dictionaty of Music. e Edition. (new entry), Cambridge.
Harvard Press, 2003
63. Tramo MJ, Koh CK, Shah GD, Braida LD Effects of bilateral auditory cortex lesions on frequency
processing and virtual pitch perception. ARO Abstracts 2004; 27:78
64. Tramo MJ Physiology, anatomy, and plasticity of the cerebral cortex in relation to musical instrument
performance. Journal of the Acoustical Society of America 2004 115; 2590
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65. Tramo MJ, Koh CK, Shah GD, Braida LD Functional role of the auditory cortex in virtual pitch
perception. Society for neuroscience Abstracts 2004
66. Tramo MJ, Cariani PA, Koh CK, Mathis N, Braida LD Neurophysiology and neuroanatomy of pitch
perception. Annals of the New York Academy of Sciences 2005; 1060:148-174.
67. Koss AD, Tramo MJ, Flaherty AW, Young AB Effect of auditory stimulation with popular music on
visuomotor integration and gait in Parkinson's disease. Neurology 2006; 67:114
68. Tramo MJ, Koh CK, Lense MD, Van Ness CM, Krishnamoorthy KS. Kagan 3, Caviness VS Effect of
auditory stimulation with vocal music on neurophysiologic responses to pain in premature infants.
Society for Neuroscience Abstracts 2006
69. Dykstra AD. Braids LD, Koh CK, Tramo MJ Intensity discrimination following bilateral lesions of
auditory cortex. Cognitive Neuroscience Society Abstracts 2007 #O93
70. Dykstra AD. Braida LD, Koh CK, Tramo MI Role of primary auditory cortex in intensity processing and
loudness perception. Music Language Mind Abstracts 2008
71. Dykstra AD, Braida LD, Koh CK, Tramo MJ Primary and secondary auditory cortex are necessary for
fine-grained intensity processing and loudness discriminaion. Society for Neuroscience Abstracts 2010
72. Dykstra AD, Koh CK, Braida LD, Tramo MJ Functional role of auditory cortex in intensity processing
and loudness perception. PLoS One (in revision)
73. Tramo MJ, Lense M. Van Ness C, Kagan J Effect of vocal music on physiological and behavioral
responses to pain and stress in premature infants. Music and Medicine (under review)
74. Tramo MI, Sema M, Koelsch S A cortical network for hierarchical processing of musical phrase
structure. Proceedings of the National Academy of Sciences 2010 (in revision)
In preparation
75. Tramo MJ, Cognitive Neuroscience of Music, 2nd Edition. London: Oxford Press 2010 (under contract)
76. Tramo MI, Cariani PA, Delgutte B Neurobiological and psychophysical foundations of Western tonal
harmony.
77. Tramo MJ, Chen K. Koh CK, Rekdahl AJ, Dykstra AD. Braida LD Activity dependent auditory acuity:
Harmony perception.
78. Tramo MJ, Li L, Gazzaniga MS Timbre discrimination and recognition by the right and left
hemispheres of split-brain patients.
80. Tramo MJ, Bonn CD, Shah GD, Koh CK, Braida LD Functional role of primary auditory cortex in
virtual pitch perception.
81. Tramo MJ, Koh CK, Braida LD Functional effects of bilateral auditory midbrain lesions
82. Tramo MJ, Rubin S, Bonn CD, Symons J Neuroaeshetics of music. Heightened emotional responses
to music in a conductor following bilateral ventromedial frontal lobe lesions.
83. Tramo MJ, Symons J, Gow D Acoustic phonetics of lyrics.
84. Tramo MJ, Dementia: Diagnosis, Pathophysiology & Treatment. NY: Oxford Press 2011 (under
contract)
B. Educational Material
. Tramo MI Sensory and perceptual systems. In: Cognitive and Behavioral Neurology: Focus on
Dementia, 1997-8 Course syllabus, Harvard Medical School Department of Continuing Medical
Education
2. Tramo MJ Psychology 987b. Music. Mind, and Brain, 1998-2006. Course syllabus. Harvard College
3. Tramo MJ, Functional organization of the cerebral cortex. In: Dementia: A Comprehensive Update,
2001-7 Course syllabus, Harvard Medical School Department of Continuing Medical Education
C. Thesis
. Tramo MJ Neural representations of acoustic information in relation to music and voice perception.
Harvard University, Cambridge, MA, Ph.D. Dissertation, 1998
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D. Nonprint Material
I. www.BrainMusic.org, The Institute for Music and Brain Science
2. www.myspace.comfhlenWithTales
PART IV. Copyrights, Performances, & Credits (Musical and Dramaticomusical)
Top of the Pop, World's Fair, New York, NY. 1965
Apotheosis of a King Who Lost His Kingdom, Script. Lyrics, & Music O1974 Tramo
Apotheosis of a King Who Lost His Kingdom, Sound Recording O1976 Tramo
Dynamite Tonight, Yale Repertory Theatre, 1975
Creating for the Musical Theatre. Yale Music School (Lehman Engel), 1975
Dramatic Technical Production, Yale Drama School & Repertory Theatre (John Hood, Walter Jones), 1975
Alice in Wonderland, Louis Barton, Yale Music School
Benefit, Men With Tales, Yale University, 1977
In Concert, Men With Tales. Yale University. 1978
Siren, Words & Music O1981 Tramo & Schiller
Living In a Fantasy, Music & Lyrics O1981 Tramo & Schiller
Leslie, Music & Lyrics O1981 Schiller & Tramo
Men With Tales, Sound Recording O1981 Frost. Schiller. Singleton. Tramo & Trombetta
Nadine, Sound Recording, Music & Lyrics O1987 Tramo
Party, Party, Sound Recording, Music & Lyrics O1987 Tramo
Saw You In My Dream, Sound Recording, Music & Lyrics O1988 Tramo
Hold Me, Music & Lyrics O1991 Tramo
Few of Many, Sound Recording. Music & Lyrics O2002 Tramo
For Your Heart, Sound Recording, Music & Lyrics 02002 Tramo
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